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prevented¡¡the¡¡passage¡¡of¡¡birds¡¡from¡¡one¡¡island¡¡to¡¡another£»¡¡so¡¡that¡¡out¡¡of¡¡at¡¡least¡¡three¡¡hundred¡¡and¡¡fifty¡¡land¡¡birds¡¡inhabiting¡¡Java¡¡and¡¡Borneo£»¡¡not¡¡more¡¡than¡¡ten¡¡have¡¡passed¡¡eastward¡¡into¡¡Celebes¡£¡¡Yet¡¡the¡¡Straits¡¡of¡¡Macassar¡¡are¡¡not¡¡nearly¡¡so¡¡wide¡¡as¡¡the¡¡Java¡¡sea£»¡¡and¡¡at¡¡least¡¡a¡¡hundred¡¡species¡¡are¡¡common¡¡to¡¡Borneo¡¡and¡¡Java¡£

I¡¡will¡¡now¡¡give¡¡two¡¡examples¡¡to¡¡show¡¡how¡¡a¡¡knowledge¡¡of¡¡the¡¡distribution¡¡of¡¡animals¡¡may¡¡reveal¡¡unsuspected¡¡facts¡¡in¡¡the¡¡past¡¡history¡¡of¡¡the¡¡earth¡£¡¡At¡¡the¡¡eastern¡¡extremity¡¡of¡¡Sumatra£»¡¡and¡¡separated¡¡from¡¡it¡¡by¡¡a¡¡strait¡¡about¡¡fifteen¡¡miles¡¡wide£»¡¡is¡¡the¡¡small¡¡rocky¡¡island¡¡of¡¡Banca£»¡¡celebrated¡¡for¡¡its¡¡tin¡¡mines¡£¡¡One¡¡of¡¡the¡¡Dutch¡¡residents¡¡there¡¡sent¡¡some¡¡collections¡¡of¡¡birds¡¡and¡¡animals¡¡to¡¡Leyden£»¡¡and¡¡among¡¡them¡¡were¡¡found¡¡several¡¡species¡¡distinct¡¡from¡¡those¡¡of¡¡the¡¡adjacent¡¡coast¡¡of¡¡Sumatra¡£¡¡One¡¡of¡¡these¡¡was¡¡a¡¡squirrel¡¡£¨Sciurus¡¡bangkanus£©£»¡¡closely¡¡allied¡¡to¡¡three¡¡other¡¡species¡¡inhabiting¡¡respectively¡¡the¡¡Malay¡¡peninsula£»¡¡Sumatra£»¡¡and¡¡Borneo£»¡¡but¡¡quite¡¡as¡¡distinct¡¡from¡¡them¡¡all¡¡as¡¡they¡¡are¡¡from¡¡each¡¡other¡£¡¡There¡¡were¡¡also¡¡two¡¡new¡¡ground¡¡thrushes¡¡of¡¡the¡¡genus¡¡Pitta£»¡¡closely¡¡allied¡¡to£»¡¡but¡¡quite¡¡distinct¡¡from£»¡¡two¡¡other¡¡species¡¡inhabiting¡¡both¡¡Sumatra¡¡and¡¡Borneo£»¡¡and¡¡which¡¡did¡¡not¡¡perceptibly¡¡differ¡¡in¡¡these¡¡large¡¡and¡¡widely¡¡separated¡¡islands¡£¡¡This¡¡is¡¡just¡¡as¡¡if¡¡the¡¡Isle¡¡of¡¡Man¡¡possessed¡¡a¡¡peculiar¡¡species¡¡of¡¡thrush¡¡and¡¡blackbird£»¡¡distinct¡¡from¡¡the¡¡birds¡¡which¡¡are¡¡common¡¡to¡¡England¡¡and¡¡Ireland¡£

These¡¡curious¡¡facts¡¡would¡¡indicate¡¡that¡¡Banca¡¡may¡¡have¡¡existed¡¡as¡¡a¡¡distinct¡¡island¡¡even¡¡longer¡¡than¡¡Sumatra¡¡and¡¡Borneo£»¡¡and¡¡there¡¡are¡¡some¡¡geological¡¡and¡¡geographical¡¡facts¡¡which¡¡render¡¡this¡¡not¡¡so¡¡improbable¡¡as¡¡it¡¡would¡¡at¡¡first¡¡seem¡¡to¡¡be¡£¡¡Although¡¡on¡¡the¡¡map¡¡Banca¡¡appears¡¡so¡¡close¡¡to¡¡Sumatra£»¡¡this¡¡does¡¡not¡¡arise¡¡from¡¡its¡¡having¡¡been¡¡recently¡¡separated¡¡from¡¡it£»¡¡for¡¡the¡¡adjacent¡¡district¡¡of¡¡Palembang¡¡is¡¡new¡¡land£»¡¡being¡¡a¡¡great¡¡alluvial¡¡swamp¡¡formed¡¡by¡¡torrents¡¡from¡¡the¡¡mountains¡¡a¡¡hundred¡¡miles¡¡distant¡£

Banca£»¡¡on¡¡the¡¡other¡¡hand£»¡¡agrees¡¡with¡¡Malacca£»¡¡Singapore£»¡¡and¡¡the¡¡intervening¡¡island¡¡of¡¡Lingen£»¡¡in¡¡being¡¡formed¡¡of¡¡granite¡¡and¡¡laterite£»¡¡and¡¡these¡¡have¡¡all¡¡most¡¡likely¡¡once¡¡formed¡¡an¡¡extension¡¡of¡¡the¡¡Malay¡¡peninsula¡£¡¡As¡¡the¡¡rivers¡¡of¡¡Borneo¡¡and¡¡Sumatra¡¡have¡¡been¡¡for¡¡ages¡¡filling¡¡up¡¡the¡¡intervening¡¡sea£»¡¡we¡¡may¡¡be¡¡sure¡¡that¡¡its¡¡depth¡¡has¡¡recently¡¡been¡¡greater£»¡¡and¡¡it¡¡is¡¡very¡¡probable¡¡that¡¡those¡¡large¡¡islands¡¡were¡¡never¡¡directly¡¡connected¡¡with¡¡each¡¡other¡¡except¡¡through¡¡the¡¡Malay¡¡peninsula¡£¡¡At¡¡that¡¡period¡¡the¡¡same¡¡species¡¡of¡¡squirrel¡¡and¡¡Pitta¡¡may¡¡have¡¡inhabited¡¡all¡¡these¡¡countries£»¡¡but¡¡when¡¡the¡¡subterranean¡¡disturbances¡¡occurred¡¡which¡¡led¡¡to¡¡the¡¡elevation¡¡of¡¡the¡¡volcanoes¡¡of¡¡Sumatra£»¡¡the¡¡small¡¡island¡¡of¡¡Banca¡¡may¡¡have¡¡been¡¡separated¡¡first£»¡¡and¡¡its¡¡productions¡¡being¡¡thus¡¡isolated¡¡might¡¡be¡¡gradually¡¡modified¡¡before¡¡the¡¡separation¡¡of¡¡the¡¡larger¡¡islands¡¡had¡¡been¡¡completed¡£

As¡¡the¡¡southern¡¡part¡¡of¡¡Sumatra¡¡extended¡¡eastward¡¡and¡¡formed¡¡the¡¡narrow¡¡straits¡¡of¡¡Banca£»¡¡many¡¡birds¡¡and¡¡insects¡¡and¡¡some¡¡Mammalia¡¡would¡¡cross¡¡from¡¡one¡¡to¡¡the¡¡other£»¡¡and¡¡thus¡¡produce¡¡a¡¡general¡¡similarity¡¡of¡¡productions£»¡¡while¡¡a¡¡few¡¡of¡¡the¡¡older¡¡inhabitants¡¡remained£»¡¡to¡¡reveal¡¡by¡¡their¡¡distinct¡¡forms£»¡¡their¡¡different¡¡origin¡£¡¡Unless¡¡we¡¡suppose¡¡some¡¡such¡¡changes¡¡in¡¡physical¡¡geography¡¡to¡¡have¡¡occurred£»¡¡the¡¡presence¡¡of¡¡peculiar¡¡species¡¡of¡¡birds¡¡and¡¡mammals¡¡in¡¡such¡¡an¡¡island¡¡as¡¡Banca¡¡is¡¡a¡¡hopeless¡¡puzzle£»¡¡and¡¡I¡¡think¡¡I¡¡have¡¡shown¡¡that¡¡the¡¡changes¡¡required¡¡are¡¡by¡¡no¡¡means¡¡so¡¡improbable¡¡as¡¡a¡¡mere¡¡glance¡¡at¡¡the¡¡map¡¡would¡¡lead¡¡us¡¡to¡¡suppose¡£

For¡¡our¡¡next¡¡example¡¡let¡¡us¡¡take¡¡the¡¡great¡¡islands¡¡of¡¡Sumatra¡¡and¡¡Java¡£¡¡These¡¡approach¡¡so¡¡closely¡¡together£»¡¡and¡¡the¡¡chain¡¡of¡¡volcanoes¡¡that¡¡runs¡¡through¡¡them¡¡gives¡¡such¡¡an¡¡air¡¡of¡¡unity¡¡to¡¡the¡¡two£»¡¡that¡¡the¡¡idea¡¡of¡¡their¡¡having¡¡been¡¡recently¡¡dissevered¡¡is¡¡immediately¡¡suggested¡£¡¡The¡¡natives¡¡of¡¡Java£»¡¡however£»¡¡go¡¡further¡¡than¡¡this£»¡¡for¡¡they¡¡actually¡¡have¡¡a¡¡tradition¡¡of¡¡the¡¡catastrophe¡¡which¡¡broke¡¡them¡¡asunder£»¡¡and¡¡fix¡¡its¡¡date¡¡at¡¡not¡¡much¡¡more¡¡than¡¡a¡¡thousand¡¡years¡¡ago¡£¡¡It¡¡becomes¡¡interesting£»¡¡therefore£»¡¡to¡¡see¡¡what¡¡support¡¡is¡¡given¡¡to¡¡this¡¡view¡¡by¡¡the¡¡comparison¡¡of¡¡their¡¡animal¡¡productions¡£

The¡¡Mammalia¡¡have¡¡not¡¡been¡¡collected¡¡with¡¡sufficient¡¡completeness¡¡in¡¡both¡¡islands¡¡to¡¡make¡¡a¡¡general¡¡comparison¡¡of¡¡much¡¡value£»¡¡and¡¡so¡¡many¡¡species¡¡have¡¡been¡¡obtained¡¡only¡¡as¡¡live¡¡specimens¡¡in¡¡captivity£»¡¡that¡¡their¡¡locality¡¡has¡¡often¡¡been¡¡erroneously¡¡given£»¡¡the¡¡island¡¡in¡¡which¡¡they¡¡were¡¡obtained¡¡being¡¡substituted¡¡for¡¡that¡¡from¡¡which¡¡they¡¡originally¡¡came¡£¡¡Taking¡¡into¡¡consideration¡¡only¡¡those¡¡whose¡¡distribution¡¡is¡¡more¡¡accurately¡¡known£»¡¡we¡¡learn¡¡that¡¡Sumatra¡¡is£»¡¡in¡¡a¡¡zoological¡¡sense£»¡¡more¡¡neatly¡¡related¡¡to¡¡Borneo¡¡than¡¡it¡¡is¡¡to¡¡Java¡£¡¡The¡¡great¡¡man¡­like¡¡apes£»¡¡the¡¡elephant£»¡¡the¡¡tapir£»¡¡and¡¡the¡¡Malay¡¡bear£»¡¡are¡¡all¡¡common¡¡to¡¡the¡¡two¡¡former¡¡countries£»¡¡while¡¡they¡¡are¡¡absent¡¡from¡¡the¡¡latter¡£¡¡Of¡¡the¡¡three¡¡long¡­tailed¡¡monkeys¡¡£¨Semnopithecus£©¡¡inhabiting¡¡Sumatra£»¡¡one¡¡extends¡¡into¡¡Borneo£»¡¡but¡¡the¡¡two¡¡species¡¡of¡¡Java¡¡are¡¡both¡¡peculiar¡¡to¡¡it¡£¡¡So¡¡also¡¡the¡¡great¡¡Malay¡¡deer¡¡£¨Rusa¡¡equina£©£»¡¡and¡¡the¡¡small¡¡Tragulus¡¡kanchil£»¡¡are¡¡common¡¡to¡¡Sumatra¡¡and¡¡Borneo£»¡¡but¡¡do¡¡not¡¡extend¡¡into¡¡Java£»¡¡where¡¡they¡¡are¡¡replaced¡¡by¡¡Tragulas¡¡javanicus¡£¡¡The¡¡tiger£»¡¡it¡¡is¡¡true£»¡¡is¡¡found¡¡in¡¡Sumatra¡¡and¡¡Java£»¡¡but¡¡not¡¡in¡¡Borneo¡£¡¡But¡¡as¡¡this¡¡animal¡¡is¡¡known¡¡to¡¡swim¡¡well£»¡¡it¡¡may¡¡have¡¡found¡¡its¡¡way¡¡across¡¡the¡¡Straits¡¡of¡¡Sunda£»¡¡or¡¡it¡¡may¡¡have¡¡inhabited¡¡Java¡¡before¡¡it¡¡was¡¡separated¡¡from¡¡the¡¡mainland£»¡¡and¡¡from¡¡some¡¡unknown¡¡cause¡¡have¡¡ceased¡¡to¡¡exist¡¡in¡¡Borneo¡£

In¡¡Ornithology¡¡there¡¡is¡¡a¡¡little¡¡uncertainty¡¡owing¡¡to¡¡the¡¡birds¡¡of¡¡Java¡¡and¡¡Sumatra¡¡being¡¡much¡¡better¡¡known¡¡than¡¡those¡¡of¡¡Borneo£»¡¡but¡¡the¡¡ancient¡¡separation¡¡of¡¡Java¡¡as¡¡an¡¡island¡¡is¡¡well¡¡exhibited¡¡by¡¡the¡¡large¡¡number¡¡of¡¡its¡¡species¡¡which¡¡are¡¡not¡¡found¡¡in¡¡any¡¡of¡¡the¡¡other¡¡islands¡£¡¡It¡¡possesses¡¡no¡¡less¡¡than¡¡seven¡¡pigeons¡¡peculiar¡¡to¡¡itself£»¡¡while¡¡Sumatra¡¡has¡¡only¡¡one¡£¡¡Of¡¡its¡¡two¡¡parrots¡¡one¡¡extends¡¡into¡¡Borneo£»¡¡but¡¡neither¡¡into¡¡Sumatra¡£¡¡Of¡¡the¡¡fifteen¡¡species¡¡of¡¡woodpeckers¡¡inhabiting¡¡Sumatra¡¡only¡¡four¡¡reach¡¡Java£»¡¡while¡¡eight¡¡of¡¡them¡¡are¡¡found¡¡in¡¡Borneo¡¡and¡¡twelve¡¡in¡¡the¡¡Malay¡¡peninsula¡£¡¡The¡¡two¡¡Trogons¡¡found¡¡in¡¡Java¡¡are¡¡peculiar¡¡to¡¡it£»¡¡while¡¡of¡¡those¡¡inhabiting¡¡Sumatra¡¡at¡¡least¡¡two¡¡extend¡¡to¡¡Malacca¡¡and¡¡one¡¡to¡¡Borneo¡£¡¡There¡¡are¡¡a¡¡very¡¡large¡¡number¡¡of¡¡birds£»¡¡such¡¡as¡¡the¡¡great¡¡Argus¡¡pheasant£»¡¡the¡¡fire¡­backed¡¡and¡¡ocellated¡¡pheasants£»¡¡the¡¡crested¡¡partridge¡¡£¨Rollulus¡¡coronatus£©£»¡¡the¡¡small¡¡Malacca¡¡parrot¡¡£¨Psittinus¡¡incertus£©£»¡¡the¡¡great¡¡helmeted¡¡hornbill¡¡£¨Buceroturus¡¡galeatus£©£»¡¡the¡¡pheasant¡¡ground¡­cuckoo¡¡£¨Carpococcyx¡¡radiatus£©£»¡¡the¡¡rose¡­crested¡¡bee¡­eater¡¡£¨Nyctiornis¡¡amicta£©£»¡¡the¡¡great¡¡gaper¡¡£¨Corydon¡¡sumatranus£©£»¡¡and¡¡the¡¡green¡­¡¡crested¡¡gaper¡¡£¨Calyptomena¡¡viridis£©£»¡¡and¡¡many¡¡others£»¡¡which¡¡are¡¡common¡¡to¡¡Malacca£»¡¡Sumatra£»¡¡and¡¡Borneo£»¡¡but¡¡are¡¡entirely¡¡absent¡¡from¡¡Java¡£¡¡On¡¡the¡¡other¡¡hand¡¡we¡¡have¡¡the¡¡peacock£»¡¡the¡¡green¡¡jungle¡¡cock£»¡¡two¡¡blue¡¡ground¡¡thrushes¡¡£¨Arrenga¡¡cyanea¡¡and¡¡Myophonus¡¡flavirostris£©£»¡¡the¡¡fine¡¡pink¡­headed¡¡dove¡¡£¨Ptilonopus¡¡porphyreus£©£»¡¡three¡¡broad¡­tailed¡¡ground¡¡pigeons¡¡£¨Macropygia£©£»¡¡and¡¡many¡¡other¡¡interesting¡¡birds£»¡¡which¡¡are¡¡found¡¡nowhere¡¡in¡¡the¡¡Archipelago¡¡out¡¡of¡¡Java¡£

Insects¡¡furnish¡¡us¡¡with¡¡similar¡¡facts¡¡wherever¡¡sufficient¡¡data¡¡are¡¡to¡¡be¡¡had£»¡¡but¡¡owing¡¡to¡¡the¡¡abundant¡¡collections¡¡that¡¡have¡¡been¡¡made¡¡in¡¡Java£»¡¡an¡¡unfair¡¡preponderance¡¡may¡¡be¡¡given¡¡to¡¡that¡¡island¡£¡¡This¡¡does¡¡not£»¡¡however£»¡¡seem¡¡to¡¡be¡¡the¡¡case¡¡with¡¡the¡¡true¡¡Papilionidae¡¡or¡¡swallow¡­tailed¡¡butterflies£»¡¡whose¡¡large¡¡size¡¡and¡¡gorgeous¡¡colouring¡¡has¡¡led¡¡to¡¡their¡¡being¡¡collected¡¡more¡¡frequently¡¡than¡¡other¡¡insects¡£¡¡Twenty¡­seven¡¡species¡¡are¡¡known¡¡from¡¡Java£»¡¡twenty¡­nine¡¡from¡¡Borneo£»¡¡and¡¡only¡¡twenty¡­one¡¡from¡¡Sumatra¡£¡¡Four¡¡are¡¡entirely¡¡confined¡¡to¡¡Java£»¡¡while¡¡only¡¡two¡¡are¡¡peculiar¡¡to¡¡Borneo¡¡and¡¡one¡¡to¡¡Sumatra¡£¡¡The¡¡isolation¡¡of¡¡Java¡¡will£»¡¡however£»¡¡be¡¡best¡¡shown¡¡by¡¡grouping¡¡the¡¡islands¡¡in¡¡pairs£»¡¡and¡¡indicating¡¡the¡¡number¡¡of¡¡species¡¡common¡¡to¡¡each¡¡pair¡£¡¡Thus£º


Borneo¡¡¡¡¡£¡¡¡£¡¡¡¡¡£¡¡¡¡¡£¡¡¡£¡¡29¡¡species¡¡Sumatra¡¡¡£¡¡¡£¡¡¡¡¡£¡¡¡¡¡£¡¡¡£¡¡21¡¡¡¡do¡£¡¡¡¡¡¡¡¡20¡¡species¡¡common¡¡to¡¡both¡¡islands¡£

Borneo¡¡¡¡¡£¡¡¡£¡¡¡¡¡£¡¡¡¡¡£¡¡¡£¡¡29¡¡¡¡do¡£¡¡¡¡¡¡Java¡¡¡£¡¡¡¡¡£¡¡¡£¡¡¡¡¡£¡¡¡¡¡£¡¡¡£¡¡27¡¡¡¡do¡£¡¡¡¡¡¡¡¡20¡¡¡¡¡¡¡¡do¡£¡¡¡¡¡¡¡¡do¡£

Sumatra¡¡¡£¡¡¡£¡¡¡¡¡£¡¡¡¡¡£¡¡¡£¡¡21¡¡¡¡do¡£¡¡Java¡¡¡£¡¡¡¡¡£¡¡¡£¡¡¡¡¡£¡¡¡¡¡£¡¡¡£¡¡27¡¡¡¡do¡£¡¡¡¡¡¡¡¡11¡¡¡¡¡¡¡¡do¡£¡¡¡¡¡¡¡¡do¡£

Making¡¡some¡¡allowance¡¡for¡¡our¡¡imperfect¡¡knowledge¡¡of¡¡the¡¡Sumatran¡¡species£»¡¡we¡¡see¡¡that¡¡Java¡¡is¡¡more¡¡isolated¡¡from¡¡the¡¡two¡¡larger¡¡islands¡¡than¡¡they¡¡are¡¡from¡¡each¡¡other£»¡¡thus¡¡entirely¡¡confirming¡¡the¡¡results¡¡given¡¡by¡¡the¡¡distribution¡¡of¡¡birds¡¡and¡¡Mammalia£»¡¡and¡¡rendering¡¡it¡¡almost¡¡certain¡¡that¡¡the¡¡last¡­named¡¡island¡¡was¡¡the¡¡first¡¡to¡¡be¡¡completely¡¡separated¡¡from¡¡the¡¡Asiatic¡¡continent£»¡¡and¡¡that¡¡the¡¡native¡¡tradition¡¡of¡¡its¡¡having¡¡been¡¡recently¡¡separated¡¡from¡¡Sumatra¡¡is¡¡entirely¡¡without¡¡foundation¡£

We¡¡are¡¡now¡¡able¡¡to¡¡trace¡¡out¡¡with¡¡some¡¡probability¡¡the¡¡course¡¡of¡¡events¡£¡¡Beginning¡¡at¡¡the¡¡time¡¡when¡¡the¡¡whole¡¡of¡¡the¡¡Java¡¡sea£»¡¡the¡¡Gulf¡¡of¡¡Siam£»¡¡and¡¡the¡¡Straits¡¡of¡¡Malacca¡¡were¡¡dry¡¡land£»¡¡forming¡¡with¡¡Borneo£»¡¡Sumatra£»¡¡and¡¡Java£»¡¡a¡¡vast¡¡southern¡¡prolongation¡¡of¡¡the¡¡Asiatic¡¡continent£»¡¡the¡¡first¡¡movement¡¡would¡¡be¡¡the¡¡sinking¡¡down¡¡of¡¡the¡¡Java¡¡sea£»¡¡and¡¡the¡¡Straits¡¡of¡¡Sunda£»¡¡consequent¡¡on¡¡the¡¡activity¡¡of¡¡the¡¡Javanese¡¡volcanoes¡¡along¡¡the¡¡southern¡¡extremity¡¡of¡¡the¡¡land£»¡¡and¡¡leading¡¡to¡¡the¡¡complete¡¡separation¡¡of¡¡that¡¡island¡£¡¡As¡¡the¡¡volcanic¡¡belt¡¡of¡¡Java¡¡and¡¡Sumatra¡¡increased¡¡in¡¡activity£»¡¡more¡¡and¡¡more¡¡of¡¡the¡¡land¡¡was¡¡submerged£»¡¡until¡¡first¡¡Borneo£»¡¡and¡¡afterwards¡¡Sumatra£»¡¡became¡¡entirely¡¡severed¡£¡¡Since¡¡the¡¡epoch¡¡of¡¡the¡¡first¡¡disturbance£»¡¡several¡¡distinct¡¡elevations¡¡and¡¡depressions¡¡may¡¡have¡¡taken¡¡pl

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