the malay archipelago-1-第33节
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n most of the islands; than we do of the plants。 It is these two classes which will supply us with most of our facts as to the geographical distribution of organized beings in this region。
The number of Mammalia known to inhabit the Indo…Malay region is very considerable; exceeding 170 species。 With the exception of the bats; none of these have any regular means of passing arms of the sea many miles in extent; and a consideration of their distribution must therefore greatly assist us in determining whether these islands have ever been connected with each other or with the continent since the epoch of existing species。
The Quadrumana or monkey tribe form one of the most characteristic features of this region。 Twenty…four distinct species are known to inhabit it; and these are distributed with tolerable uniformity over the islands; nine being found in Java; ten in the Malay peninsula; eleven in Sumatra; and thirteen in Borneo。 The great man…like Orangutans are found only in Sumatra and Borneo; the curious Siamang (next to them in size) in Sumatra and Malacca; the long…nosed monkey only in Borneo; while every island has representatives of the Gibbons or long…armed apes; and of monkeys。 The lemur…like animals; Nycticebus; Tarsius; and Galeopithecus; are found on all the islands。
Seven species found on the Malay peninsula extend also into Sumatra; four into Borneo; and three into Java; while two range into Siam and Burma; and one into North India。 With the exception of the Orangutan; the Siamang; the Tarsius spectrum; and the Galeopithecus; all the Malayan genera of Quadrumana are represented in India by closely allied species; although; owing to the limited range of most of these animals; so few are absolutely identical。
Of Carnivora; thirty…three species are known from the Indo…Malay region; of which about eight are found also in Burma and India。 Among these are the tiger; leopard; a tiger…cat; civet; and otter; while out of the twenty genera of Malayan Carnivora; thirteen are represented in India by more or less closely allied species。 As an example; the Malayan bear is represented in North India by the Tibetan bear; both of which may be seen alive at the Zoological Society's Gardens。
The hoofed animals are twenty…two in number; of which about seven extend into Burmahand India。 All the deer are of peculiar species; except two; which range from Malacca into India。 Of the cattle; one Indian species reaches Malacca; while the Bos sondiacus of Java and Borneo is also found in Siam and Burma。 A goat…like animal is found in Sumatra which has its representative in India; while the two…horned rhinoceros of Sumatra and the single…horned species of Java; long supposed to be peculiar to these islands; are now both ascertained to exist in Burma; Pegu; and Moulmein。 The elephant of Sumatra; Borneo; and Malacca is now considered to be identical with that of Ceylon and India。
In all other groups of Mammalia the same general phenomena recur。 A few species are identical with those of India。 A much larger number are closely allied or representative forms; while there are always a small number of peculiar genera; consisting of animals unlike those found in any other part of the world。 There are about fifty bats; of which less than one…fourth are Indian species; thirty…four Rodents (squirrels; rats; &c。); of which six or eight only are Indian; and ten Insectivora; with one exception peculiar to the Malay region。 The squirrels are very abundant and characteristic; only two species out of twenty…five extending into Siam and Burma。 The Tupaias are curious insect…eaters; which closely resemble squirrels; and are almost confined to the Malay islands; as;are the small feather…tailed Ptilocerus lowii of Borneo; and the curious long…snouted and naked…tailed Gymnurus rafllesii。
As the Malay peninsula is a part of the continent of Asia; the question of the former union of the islands to the mainland will be best elucidated by studying the species which are found in the former district; and also in some of the islands。 Now; if we entirely leave out of consideration the bats; which have the power of flight; there are still forty…eight species of mammals common to the Malay peninsula and the three large islands。 Among these are seven Quadrumana (apes; monkeys; and lemurs); animals who pass their whole existence in forests; who never swim; and who would be quite unable to traverse a single mile of sea; nineteen Carnivora; some of which no doubt might cross by swimming; but we cannot suppose so large a number to have passed in this way across a strait which; except at one point; is from thirty to fifty miles wide; and five hoofed animals; including the Tapir; two species of rhinoceros; and an elephant。 Besides these there are thirteen Rodents and four Insectivora; including a shrew…mouse and six squirrels; whose unaided passage over twenty miles of sea is even more inconceivable than that of the larger animals。
But when we come to the cases of the same species inhabiting two of the more widely separated islands; the difficulty is much increased。 Borneo is distant nearly 150 miles from Biliton; which is about fifty miles from Banca; and this fifteen from Sumatra; yet there are no less than thirty…six species of mammals common to Borneo and Sumatra。 Java again is more than 250 miles from Borneo; yet these two islands have twenty…two species in common; including monkeys; lemurs; wild oxen; squirrels and shrews。 These facts seem to render it absolutely certain that there has been at some former period a connection between all these islands and the mainland; and the fact that most of the animals common to two or more of then; show little or no variation; but are often absolutely identical; indicates that the separation must have been recent in a geological sense; that is; not earlier than the Newer Pliocene epoch; at which time land animals began to assimilate closely with those now existing。
Even the bats furnish an additional argument; if one were needed; to show that the islands could not have been peopled from each other and from the continent without some former connection。 For if such had been the mode of stocking them with animals; it is quite certain that creatures which can fly long distances would be the first to spread from island to island; and thus produce an almost perfect uniformity of species over the whole region。 But no such uniformity exists; and the bats of each island are almost; if not quite; as distinct as the other mammals。 For example; sixteen species are known in Borneo; and of these ten are found in Java and five in Sumatra; a proportion about the same as that of the Rodents; which have no direct means of migration。 We learn from this fact; that the seas which separate the islands from each other are wide enough to prevent the passage even of flying animals; and that we must look to the same causes as having led to the present distribution of both groups。 The only sufficient cause we can imagine is the former connection of all the islands with the continent; and such a change is in perfect harmony with what we know of the earth's past history; and is rendered probable by the remarkable fact that a rise of only three hundred feet would convert the wide seas that separate them into an immense winding valley or plain about three hundred miles wide and twelve hundred long。 It may; perhaps; be thought that birds which possess the power of flight in so pre…eminent a degree; would not be limited in their range by arms of the sea; and would thus afford few indications of the former union or separation of the islands they inhabit。 This; however; is not the case。 A very large number of birds appear to be as strictly limited by watery barriers as are quadrupeds; and as they have been so much more attentively collected; we have more complete materials to work upon; and are able to deduce from them still more definite and satisfactory results。 Some groups; however; such as the aquatic birds; the waders; and the birds of prey; are great wanderers; other groups are little known except to ornithologists。 I shall therefore refer chiefly to a few of the best known and most remarkable families of birds as a sample of the conclusions furnished by the entire class。
The birds of the Indo…Malay region have a close resemblance to those of India; for though a very large proportion of the species are quite distinct; there are only about fifteen peculiar genera; and not a single family group confined to the former district。 If; however; we compare the islands with the Burmese; Siamese; and Malayan countries; we shall find still less difference; and shall be convinced that all are closely united by the bond of a former union。 In such well…known families as the woodpeckers; parrots; trogons; barbets; kingfishers; pigeons; and pheasants; we find some identical species spreading over all India; and as far as Java and Borneo; while a very large proportion are common to Sumatra and the Malay peninsula。
The force of these facts can only be appreciated when we come to treat the islands of the Austro…Malay region; and show how similar barriers have entirely prevented the passage of birds from one island to another; so that out of at least three hundred and